It looked as though Dele Alli's penalty would seal an undeserved three points for Spurs to send them into Sunday's north London derby on the back of a win.
It was substitute Karamoko had other ideas and sent a fine strike beyond Joe Hart from 20 yards to make sure the spoils were shared in a strange but memorable Group J clash.
Tottenham went behind to Peter Michorl's excellent 42nd-minute opener.
However, Gareth Bale levelled from the spot with the last kick of the first half and Son Heung-min put Spurs ahead soon after the restart.
Johannes Eggestein levelled with six minutes remaining but there was still time for Alli - on the field for just four minutes - to make his mark from 12 yards after Steven Bergwijn was brought down by Philipp Wiesinger.
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